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The problem of whether haplodiploidy is responsible for the frequent evolution of eusociality in the Hymenoptera remains unresolved. The little-known "protected invasion hypothesis" posits that because a male will transmit a new allele for alloparental care to all his daughters under haplodiploidy, such an allele has a higher probability of spreading to fixation under haplodiploidy than under diploidy. This mechanism is investigated using the mating system and lifecycles ancestral to eusocial lineages. It is shown that although haplodipl
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