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The majority of freshwater ecosystems worldwide suffer from eutrophication, particularly because of agriculture-derived nutrient sources. In the European Union, a discrepancy exists between the scale of regulatory assessment and the size of research catchments. The Water Framework Directive sets water quality objectives at the mesoscale (50-500 km2), a scale at which both hillslope and in-stream processes influence carbon (C), nitrogen (N) and phosphorus (P) dynamics. Conversely, research catchments focus on headwaters to investigate h
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