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Changes in the local granular microstructure caused by NBs were favorable for the porous structure of granules to avoid granular disintegration and flotation caused by the excessive secretion of extracellular polymeric substances blocking gas channels. The formation and stability of NBs penetrating the cell clusters played a crucial role in the formation and stability of nanopores around or inside the cell clusters, further providing a basis for the formation of high-porosity structures and efficient mass transfer of AnGS.In this study, t
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