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Root hair (RH) size has vital physiological implications, since it influences the surface area of the root and thus the ability of the plant to absorb water and nutrients from the soil. Arabidopsis thaliana ROOT HAIR DEFECTIVE 6-LIKE 4 (RSL4), a bHLH transcription factor, controls the expression of hundreds of RH genes and RSL4 expression itself can trigger ectopic RH growth. Recent studies reveal an autocrine mechanism governing plant RH cell growth in which the extracellular peptide RAPID ALKALINIZATION FACTOR 1 (RALF1) and receptor F
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