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As is the case today, both climate variability and population density influenced human behavioural change in the past. The mechanisms underpinning later Pleistocene human behavioural evolution, however, remain contested. Many complex behaviours evolved in Africa, but early evidence for these behaviours varies both spatially and temporally. Scientists have not been able to explain this flickering pattern, which is present even in sites and regions clearly occupied by Homo sapiens. To explore this pattern, here the presence and frequency o
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