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Making use of allele frequency directional difference and thickness (AFDDD) mapping strategy, one region on chromosome 4 and another on chromosome 6 had been identified become putatively connected with large acidity, and were called Ma6 and Ma4, correspondingly. Trait association analysis of DNA markers independently developed from the Ma6 and Ma4 areas confirmed the mapping of Ma6 and Ma4. In the back ground of MaMa, 20.6% of acidity variation could be explained by Ma6, 28.5% by Ma4, and 50.7% by
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