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Botanical nastic systems demonstrate non-directional structural responses to stimuli such as pressure, light, chemicals or temperature; hygronasty refers to systems that respond specifically to moisture. Many seed dispersal mechanisms such as wheat awns, legume pods, spruce and pinecones fall within this classification. The variety of behaviours varies greatly from opening and closing to self-digging, but the mechanism is based on differential hygroscopic swelling between two adjacent areas of tissue. We describe the application of hygr
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