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PRACTITIONER POINTS Evaluated alone τAN exhibits no statistical effect on effluent phosphorus in an EBPR configuration. Increased PHA synthesis, associated with increased VFAs and/or extended τAN, enhances aerobic phosphorus removal. PHA synthesis normalized to VFA loading increased with τAN , suggesting fermentation in the EBPR anaerobic zone. Aerobic phosphorus uptake increases linearly with anaerobic phosphorus release, with the slope exceeding unity. Increased VFAs can be substituted for shorter anaerobic HRTs, and vice versa, to enha
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